In a fascinating talk, Chad Watkins reported the discovery of a surprising frequency of homeotic mutations along the anterior-posterior axis and implicated the high rate of transposons in the Hox D cluster as a possible cause. Watkins reported that 27% of DNA in the HoxD cluster is comprised of transposons, much higher than in other vertebrates. Moreover, Hox D gene expression is shifted posteriorly. For example, Hox D10 normally should express at anterior boundary of hip, but in A. carolinensis it is all the way down at the tip of the tail. Further, in later stage embryo, ectopic expression of D13 occurs in the hand and feet, which is novel in vertebrates. Could this be related to ecomorph differentiation in limb elements?