A while back we had a post featuring a photo of Anolis limifrons in full battle mode, tongues sticking out. Check out the comments on that post for discussion of the prevalence of this behavior. One commenter said that he’d seen it in A. cupreus, and now here’s visual proof.
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Peter Mudde
Anolis aquaticus sticks out its tongue as well during some displays. The male is a hybrid with parents from near Dominical and San Vito in Costa Rica.